The Deathwing Astartes must clear out these ancient vessels of their xenos infestation to succeed at various objectives assigned to them by the Dark Angels' secretive leadership, called the Inner Circle. The primary antagonist of the game are the species of Tyranids known as Genestealers, who infest large, ancient conglomerate starships called Space Hulks. An example of one such psychic ability that Librarians in the game can unleash is Hellfire, which sends psychic fire coursing towards the enemy. A Librarian can use the traditional weaponry of the Space Marines, such as the Bolter and a Power Sword, but also their own psychic might against their foes. Librarians, unlike other Space Marines, can use the psychic power of the Warp, also known as the Immaterium. In Space Hulk: Deathwing the single player campaign puts the player in the shoes of a Librarian of the Dark Angels Chapter's elite 1st Company of Space Marines, known as the Deathwing, as they carry out a series of missions for the Chapter while wearing Tactical Dreadnought Armour.

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